Etan Markus

Professor & Associate Department Head

of Graduate Studies

Degree: Ph.D., Year, University/College

Ph.D., 1989   University of Toronto, Toronto

Webpages: Markus Lab

Precollege Program

UCONN Brain&Behavior@TAU: TelAviv Summer Psychology/Neuroscience Program

Markus PSYC2209 Website – Learning & memory: From Brain to Behavior

Research Interests: Brain basis of behavior, focusing on rats. Specifically:

Memory formation


Hippocampal system


Social Interactions

Research Synopsis: The broad goal is understanding how an experience gets “turned into” a memory. The lab focuses on how neurons process and represent information in a specific brain system, the hippocampus. The hippocampus is important for episodic memory and knowing your location while navigating through the environment. Techniques used include training rats on an array of behavioral tasks (mazes), brain inactivation, and monitoring the activity of individual cells during behavior.
Courses: PSYC 2500 Learning

PSYC 3250w Animal Behavior & Learning Lab

PSYC 5351: Neural Foundations of Learning and Memory

PSYC 5285: Neurobiology of Aging: Changes in Cognitive Processes

Recent publications: See publications at
Representative Publications: Jacobson TK, Schmidt B, Hinman JR, Escabí MA, Markus EJ. (2015). Age-related decrease in theta and gamma coherence across dorsal ca1 pyramidale and radiatum layers. Hippocampus 25(11):1327-35


Schmidt B (g), Hinman JR, Jacobson TK (g), Szkudlarek E (u), Argraves M (u), Escabí MA, Markus EJ (2013). Dissociation between Dorsal and Ventral Hippocampal Theta Oscillations during Decision-Making. J. Neuroscience 33(14):6212-24


Jacobson TK, Howe MD, Schmidt B, Hinman JR, Escabi M, Markus EJ (2013). Hippocampal theta, gamma, and theta-gamma coupling: Effects of aging, environmental change, and cholinergic activation. J. Neurophysiology 109(7):1852-65


Schmidt B, Papale A, Redish AD & Markus EJ (2013). Conflict between Place and Response Navigation Strategies: Effects on Vicarious Trial and Error (VTE) Behaviors. Learning & Memory 15;20(3):130-8


Schmidt B, Marrone D, and Markus EJ (2011).   Disambiguating the similar: The dentate gyrus and pattern separation. Behavioral Brain Research. 226(1):56-65

Honors & Awards: 2016 AAUP Teaching Excellence Career Award

2016 CLAS Excellence in Teaching Award

2016 OURs Faculty Mentorship Excellence Award


Etan Markus
Contact Information
Mailing AddressUnit 1020
Office LocationBousfield 100